# A gradient of the HD-Zip regulator Woolly regulates multicellular trichome morphogenesis in tomato

*PRI Rank #17 · Topics and Trends in Most Cited Plant Physiology and Cultivation Studies Papers, Class of 2026*

*Canonical URL: https://pri.pepkio.com/top-papers/plant-physiology-and-cultivation-studies/2026/rank-17*

| Field | Value |
| --- | --- |
| Rank | #17 |
| 18m citations | 27 |
| Journal | The Plant Cell |
| Year | 2024 |
| DOI | 10.1093/plcell/koae077 |
| Corresponding authors | Shuang Wu |
| Institution | Fujian Agriculture and Forestry University, China |

**Ranking page:** [Topics and Trends in Most Cited Plant Physiology and Cultivation Studies Papers, Class of 2026](https://pri.pepkio.com/top-papers/plant-physiology-and-cultivation-studies/2026)

**Paper link:** [10.1093/plcell/koae077](https://doi.org/10.1093/plcell/koae077)

## Topics

HD-Zip IV · Woolly (Wo) · multicellular trichomes · tomato · morphogenesis · gradient patterning · spatially polarized cell division · cell expansion · TCP transcription factor · SlBRANCHED2a (SlBRC2a) · endoreduplication · negative feedback loop · L1 epidermal cell layer · apical-basal patterning · cell division to cell expansion transition

## Cite this ranking

```
Pepkio Research Index (PRI). Topics and Trends in Most Cited Plant Physiology and Cultivation Studies Papers, Class of 2026. https://pri.pepkio.com/top-papers/plant-physiology-and-cultivation-studies/2026. Accessed 2026-07-15.

Zheng Su, Tinsley Li, Thematic Shifts in Early-High-Impact Cancer Genomics and Diagnostics Research: A Bibliometric and Semantic Analysis. bioRxiv 2026.07.04.736459; doi: https://doi.org/10.64898/2026.07.04.736459
```