# Oxygen-evolving photosystem II structures during S1–S2–S3 transitions

*PRI Rank #5 · Topics and Trends in Most Cited Photosynthetic Processes and Mechanisms Papers, Class of 2026*

*Canonical URL: https://pri.pepkio.com/top-papers/photosynthetic-processes-and-mechanisms/2026/rank-5*

| Field | Value |
| --- | --- |
| Rank | #5 |
| 18m citations | 57 |
| Journal | Nature |
| Year | 2024 |
| DOI | 10.1038/s41586-023-06987-5 |
| Corresponding authors | Michihiro Suga, Jian‐Ren Shen |
| Institution | Okayama University, Japan |

**Ranking page:** [Topics and Trends in Most Cited Photosynthetic Processes and Mechanisms Papers, Class of 2026](https://pri.pepkio.com/top-papers/photosynthetic-processes-and-mechanisms/2026)

**Paper link:** [10.1038/s41586-023-06987-5](https://doi.org/10.1038/s41586-023-06987-5)

## Topics

Photosystem II · Kok cycle · Mn4CaO5 cluster · Water oxidation · Oxygen evolution · Serial femtosecond crystallography · YZ tyrosine · P680 reaction center · D1-E189 · Ca2+ ion · O6 oxygen intermediate · O1 channel · O4 channel · Cl-1 channel · substrate water delivery · Proton-coupled electron transfer · nanosecond to millisecond dynamics · flash illumination · Oxygen evolution

## Cite this ranking

```
Pepkio Research Index (PRI). Topics and Trends in Most Cited Photosynthetic Processes and Mechanisms Papers, Class of 2026. https://pri.pepkio.com/top-papers/photosynthetic-processes-and-mechanisms/2026. Accessed 2026-07-21.

Zheng Su, Tinsley Li, Thematic Shifts in Early-High-Impact Cancer Genomics and Diagnostics Research: A Bibliometric and Semantic Analysis. bioRxiv 2026.07.04.736459; doi: https://doi.org/10.64898/2026.07.04.736459
```